ethologist

1. The theory of memetics is first proposed by Richard Dawkins, a British ethologist and evolutionary biologist in Oxford University, in his best seller book The Selfish Gene in 1976.
   1976年，英国牛津大学著名动物学家和行为生态学家RichardDawkins在他出版的畅销书《自私的基因》中提出模因论的基本思想。

noun

1. a zoologist who studies the behavior of animals in their natural habitats